Premier League clubs have spent £394.8m on new signings during the winter transfer window.

The outlay represents more than a three-fold increase on last year, when Premier League clubs spent only £116.2m after the Premier League's Profit and Sustainability Rules (PSR) slashed purchasing power.

Clubs recouped £130.7m on player sales, equating to a net spend of £264.1m - the second-highest winter total on record.

Spending during the window was largely incremental, with notable spikes on January 23 - when Manchester City splashed £63.2m on Eintracht Frankfurt forward Omar Marmoush - and Deadline Day.

Who spent the most?

Eleven clubs signed players for disclosed fees, with Manchester City shelling out a league-topping £176.1m on Marmoush, Nicolas Gonzalez (£50m from Porto), Abdukodir Khusanov (£33.5m from Lens) and Vitor Reis (£29.4m from Palmeiras).

The Cityzens also snapped up Juma Bah and Christian McFarlane for undisclosed fees from Real Valladolid and New York City, respectively.

Wolves splurged £42.6m on Emmanuel Agbadou (£16.6m from Reims), Marshall Munetsi (£16m from Reims) and Nasser Djiga (£10m from Red Star Belgrade), while Brighton spent £40m on Stefanos Tzimas (£20m from Nuremberg), Diego Gomez (£11m from Inter Miami) and Eiran Cashin (£9m from Derby).

Meanwhile, Manchester United (£29.6m), Aston Villa (£26m), Ipswich (£24m), Bournemouth (£18.1m), Spurs (£12.5m), Crystal Palace, Chelsea (both £12m) and Leicester (£1.9m) also recruited players for registered fees.

Villa still registered a £68m profit during the window after selling Jhon Duran to Al Nassr (£64m), Jaden Philogene to Ipswich (£20m) and Diego Carlos to Fenerbahce (£10m) - while spending only a quarter of that income on Donyell Malen from Borussia Dortmund (£20m) and Andres Garcia from Levante (£6m).

Three other clubs recorded profit, including Newcastle (£11m profit), Leicester (£8.1m profit) and Chelsea (£500,000 profit).

Which players cost the most?

Marmoush's £63.2m deal was the most expensive signing during the window, while new team-mates Gonzalez, Khusanov and Reis also appear in the top five - with Patrick Dorgu in fourth spot after his £29.6m switch from Lecce to Manchester United.

In terms of outgoings, Villa duo Duran and Philogene filled the club's coffers with a cool, combined £84m alone, while another four players departed for eight-figure sums.